1Woolcock Institute of Medical Research, The University of Sydney, Glebe, Australia
2Sydney Medical School, The University of Sydney, Sydney, Australia
3Dept of Pulmonary Diseases, University of Groningen, University Medical Center Groningen, Groningen, The Netherlands
4GRIAC (Groningen Research Institute for Asthma and COPD), University of Groningen, University Medical Center Groningen, Groningen, The Netherlands
5Dept of Pathology and Medical Biology, University of Groningen, University Medical Center Groningen, Groningen, The Netherlands